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Lignin has a highly-branched and amorphous chemical structure that varies considerably with biomass species and isolation techniques (Figure 1), including p -coumaryl, coniferyl, and sinapyl alcohols ( FIGURE 1 The radical coupling of these monolignols (Figure 2) results in the formation of different inner-unit linkages, mainly ether bonds (e.g., -O-4, -O-4, and 4-O-5) and C-C bonds (e.g., -1, -5, 55 and -) (Wang et al., 2016)

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